Tectiphiala ferox
Tectiphiala (tehk-tih-fee-AHL-lah) ferox (FEH-rohks) | |||||||
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Mauritius. Photo-(Société Palmophile Francophone). | |||||||
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Contents
Habitat and Distribution
Endemic to Mauritius. Tectiphiala grows in relative scrub, in mostly wet, more-or-less acid situations at elevations of about 570–650 m above sea level.Description
Moderate, solitary or clustered, spiny, pleonanthic, monoecious palm. Stem erect, bearing persistent leaf bases basally, distally free of leaf bases, ringed with leaf scars and abundant long spines with bulbous bases. Leaves pinnate, neatly abscising in mature individuals; sheaths tubular, forming a crownshaft, bearing an untidy ligule, and very densely covered in spines of varying length and abundant dark hairs; petiole rather short, adaxially with short spines, abaxially hairy; rachis bearing stiff hairs on both surfaces or on the adaxial surface alone; leaflets single-fold, very coriaceous, acute, arranged in close or distant fascicles, and fanned within the groups, adaxially glabrous, abaxially with a very dense covering of white scales, the midnerve bearing scattered ramenta, transverse veinlets obscure. Inflorescences solitary, infrafoliar, erect in bud, becoming ± pendulous, branching to 1 order, protandrous; peduncle covered in short spines at the base, above the insertion of the peduncular bract bearing a variety of short to very long spines; prophyll inserted just above the base of the peduncle, tubular, 2-keeled, completely enclosing the inflorescence in bud, splitting along the ventral midline and abscising, densely covered in stiff dark hairs; peduncular bract 1, inserted just above the prophyll, similarly hairy, abscising before anthesis; rachis scarcely evident; rachillae 3–5, congested at the apex of the peduncle, straight or flexuous, flattened and spiny at the base; rachilla bracts arranged in ca. 6 vertical rows throughout most of the rachilla length, prominent, approximate, projecting, saucer-like, rounded, each subtending a triad except at the very tip of the rachilla where subtending solitary or paired staminate flowers; bracteoles surrounding the pistillate flower unequal, one much larger than the other. Staminate flowers asymmetrical, acute, very briefly stalked, obscuring the rachilla bracts; sepals 3, often unequal, acute, briefly connate at the base; petals 3, distinct, strongly nerved when dry, angled, acute, valvate; stamens 6(–7), ± equalling the petals, filaments ± cylindrical, ± twisted and coiled, erect at the tip, anthers dorsifixed, briefly bifid at the tip, deeply bifid at the base, latrorse; pistillode usually apparent, 1/2 as long as the stamens, trifid or oblique. Pollen ellipsoidal symmetric to asymmetric; aperture a distal sulcus; ectexine tectate, coarsely perforate and/or rugulate, aperture margin similar or slightly finer; infratectum columellate; longest axis 42–66 µm [1/1]. Pistillate flowers in bud ± obscured by the staminate flowers; sepals 3, distinct, broadly imbricate, ± acute; petals 3, distinct, scarcely exceeding the sepals, broadly imbricate with briefly valvate tips; staminodes 6(–7), small, tooth-like or linear; gynoecium ovoid, unilocular, uniovulate, the stigmas not prominent, ovule large, pendulous, probably hemianatropous. Fruit asymmetrically ovoid, dark blue-black, with apical stigmatic remains; epicarp smooth, underlain by longitudinal sclereids over a layer of tannin cells, endocarp thin with round basal operculum. Seed attached by an elongate elliptical hilum, raphe branches anastomosing, endosperm homogeneous; embryo basal. Germination and eophyll not known. Cytology not studied. (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008) Editing by edric.
Culture
Cold Hardiness Zone: 10a
Comments and Curiosities
Etymology: The late Harold Moore Jr. named this palm Tectiphiala from the Latin tectus (covered, concealed) and phiala (a flat bottomed drinking vessel, saucer, bowl) with reference to the side view of the flowers, which are at first obscured by the overlapping buds.
Uses: The palm heart is edible.
Conservation: Critically Endangered - (IUCN Red List 1998) Regeneration is hampered by the invasion of alien weeds and by seed predation. The species remains in upland areas, occurring between 570 and 650 m.
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Moderate solitary very spiny pinnate-leaved palms, native to Mauritius, with conspicuous crownshafts and distinctive cup-like pits on the rachillae. Tectiphiala is distinguished from Acanthophoenix by the remarkable saucer-like bracts that subtend the triads of flowers. (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008) Discovered in 1969 by Marc D'Unienville, who was trying to locate seed of Acanthophoenix for his friend Jean - Alain Laloutte, he brought sample material of T. ferox to Mr.Laloutte, who recognised this as different to any other palm found on Mauritius. Field studies of T. ferox placed it in the Oncosperma alliance of arecoid palms, although studies of the pollen indicated that it was unlike pollen of the Oncosperma type and showed more resemblance to the pollen of Jessenia and Scheelea. This heavily armoured monoecious palm grows at a height of 500-650 metres above sea level, at two localities on the central plateau, namely: Crown Land Declerc near Mare Longue and Bois Sec which are protected reserves and where the last herds of Java deer are to be found. The central plateau receives an annual rainfall of 5000 mm, making access to these localities extremely difficult, as these palms grow in very marshy conditions. The entire natural occurring specimens number 28. The three palms visited only had single stems although literature cites T. ferox as being cespitose with mostly two stems, one larger than the other, covered with stiff grey-black spines 150 mm long. The spathe is up to 300 mm long and is densely spiny. The orange-yellow flowers are arranged in groups of two females and one male, in six vertical rows throughout the rachillae. The ovoid fruits are blue-black, 11 mm long by 6 mm in diameter and have the scent of ripe blue berries. No seeds were found in the vicinity of the adult plants but no doubt these could have been scattered by the birds feeding on the fruits. An inspection of the soil revealed this to be heavy clay, rich in Magnesium and more or less acidic. It would appear that the requirements of Tectiphiala ferox are abnormally specific and cultivation extremly difficult, although I have heard that the Durban Botanic Garden has a seedling. It would be wonderful to see this palm introduced into this garden. |
External Links
- Glossary of Palm Terms
- MODERN BOTANICAL LATIN
- "Just To Be Clear"
- http://www.palmtalk.org/forum/index.php?/topic/14569-tectiphiala-ferox/
- http://www.palmtalk.org/forum/index.php?/topic/40334-tectiphiala-ferox-in-mauritius/
- https://stampsofmauritius.org/2018/01/07/flora-and-fauna-of-mauritius-part-i-endemic-and-indigenous-plants/
References
Phonetic spelling of Latin names by edric.
Special thanks to Geoff Stein, (Palmbob) for his hundreds of photos.
Special thanks to Palmweb.org, Dr. John Dransfield, Dr. Bill Baker & team, for their volumes of information and photos.
Glossary of Palm Terms; Based on the glossary in Dransfield, J., N.W. Uhl, C.B. Asmussen-Lange, W.J. Baker, M.M. Harley & C.E. Lewis. 2008. Genera Palmarum - Evolution and Classification of the Palms. Royal Botanic Gardens, Kew. All images copyright of the artists and photographers (see images for credits).
J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008
Many Special Thanks to Ed Vaile for his long hours of tireless editing and numerous contributions.